Andrejeva E.N. & Borovichev E.A. New liverwort records from Kazakhstan. 1. // Arctoa. 2011. Vol. 20. P. 266.

South Kazachstan, right bank of the Syr-Dar‘ya River, wet bottom of the ditch, on calcareous fine soil, with female receptacle and antheridia, 22.05.1940, Vel‘tischev.

South Kazakhstan, right bank of the Syr-Dar‘ya River, wet bottom of the temporary ditch, on calcareous fine soil, with female receptacle and antheridia, 22.05.1940, Vel‘tischev.